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| HYMENOPTERA, Signiphoridae (Walker 1840) = (Thysanidae) - (Chalcidoidea) --  <Images> & <Juveniles>             Signiphoridae                Signiphoridae. (Thysanidae) 
  -- These are small, thick-bodied chalcids that attack
  scale insects, whiteflies, and other Homoptera or are hyperparasitoids of the
  chalcids that attack Homoptera.            
  Signiphoridae range in size from 0.22 to 1.7 mm.  Their color varies from black to salmon
  pink or white, but they are not metallic. 
  Sculpturing of the cuticleis is sparce compared to families
  Chalcididae and Eurytomidae..            
  The principal characteristics are a sessile Metasoma (thread-waist absent),
  the propodeum has a medium triangular zone, the club of the antennae are long
  and  unsegmented and they are preceded
  by 1-4 ring-like annelli.  The wings
  are fringed with medium to long setae, and there are short post-marginal and
  stigmal veins. Setae are very sparce on the wing membrane.             
  The genera Chartocerus and Thysanus are cosmopolitan,
  while Clytina has been reported only from Eastern Europe.  The Neotropical genus Signiphora
  has more than half of the known species.             Most species have been recovered
  from scale insects, aphids, mealybugs, psyllids and chloropid and drosophilid
  predatory flies of scale insects (Woolley & Hanson, 2006). They function
  either as parasitoids or hyperparasitoids. While the parasitoids are useful
  in biological control, the hyperparasitoids are considered harmful (Sullivan,
  1987).   = = = =
  = = = = = = = = = = =   
 References:   Please refer to 
  <biology.ref.htm>, [Additional references may be
  found at:  MELVYL Library]   Clausen, C.
  P.  1924. 
  Univ. Calif. Publ. Tech. Bull. 3: 
  253-88.   Girault, A. A. 
  1913.  A systematic monograph
  of the chalcidoid Hymenoptera of the family Signiphorinae.  Proc. U. S.
  Nat. Mus. 45:  189-233.   Hayat, M. & B. R. Subba-Rao.  1988. 
  Family Signiphoridae.  In:  B. R. Subba-Rao, & M. Hayat (eds.), Oriental Insects, Vol.
  19.  Association For The Study of
  Oriental. Insects, Gainesville, FL.  329
  p.   Prinsloo, G. L.  1980.  An illustrated
  guide to the families of African Chalcidoidea (Insecta: Hymenoptera).  Rep. So. Africa, Dept. Agr. &
  Fisheries Sci. Bull. 395.  66 p.   Quezada, J.
  R.  1967. 
  Master of Science Thesis, Department of Entomology, University of
  California, Riverside.  98 p.   Quezada,
  J., R., P. DeBach & D. Rosen.  1973.  Biological and taxonomic studies of Signiphora borinquensis new species (Hymenoptera: Signiphoridae), a
  primary parasite of diaspine scales.  Hilgardia 41(18): 
  543-604.   Rosanov,
  I.  1965. 
  Review of the genera of parasitic Hymenoptera of the family
  Signiphoridae (Hymenoptera, Chalcidoidea). 
  Rev. d'Ent. U.S.S.R. 44: 
  508-15, 866-84.   Subba Rao, B. R.  1974.  The genera of
  Signiphoridae (Hymenoptera) with descriptions of a new genus.  Bull. Ent. Res. 64:  525-31.   Sullivan,
  D. J.  1987. Insect
  hyperparasitism. Annual Review of Entomology 32:49-70.   Trjapitcyn, V. A.  1978/1987.  Family
  Signiphoridae (Thysanidae) (Signiphorids). 
  In:  G. S. Medvedev (ed.) 1987, Keys to the
  Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad. Nauk.,
  Zool. Inst., Leningrad, SSSR. (trans. fr. Russian, Amerind. Publ. Co.,
  Pvt. Ltd., New Delhi).  1341 p.   Woolley,
  J.B.  1988.  Phylogeny and classification of the Signiphoridae (Hymenoptera:
  Chalcidoidea). Systematic Entomology 13:465-501. DOI.    Woolley, J. B.
  & P. E. Hanson.  2006. Familia Signiphoridae. In: Hanson, P.
  E. & Gauld, I. D. (Eds.) Hymenoptera de la Región Neotropical. Memoirs of the American Entomological Institute 77:422-425.   Yoshimoto,
  C. M.  1984. 
  The Insects and Arachnids of Canada. Part 12.  The Families and Subfamilies of Canadian
  Chalcidoid Wasps, Hymenoptera: Chalcidoidea. 
  Biosystematics Res. Inst., Ottawa, Ontario, Res. Br. Agr. Canada Publ. 1760. 
  149 p.     |